Mutualisms

Mutualisms are ecological interactions between two species in which
both benefit. Many mutualisms involve species living closely together
(symbiosis); a species may be so dependent that it cannot live without
its mutualistic partner (
obligate mutualism). In other cases, a species can interact mutualistically with more than one partner (
diffuse mutualism) or even live without its partner(s) under certain conditions (
facultative mutualism).
Although all species involved in a mutualistic relationship contribute
to the partnership, we still expect each species to be "selfish" and to
evolve traits that provide the maximum possible fitness benefit while
minimizing cost.
1.
Trophic mutualisms are interactions in which both species receive a benefit of resources. Organisms require both nutrients and
energy to survive. In many trophic mutualisms, a plant provides energy from
photosynthesis
to a partner species. The partner, often a microbe, then provides the
plant with increased access to nutrients. Examples include
lichens (a symbiosis between a fungus and an alga), as well as associations between mycorrhizal fungi and plants, between
Rhizobium bacteria and legumes, and between corals and
zooxanthellae.
Other trophic partnerships involve animals. Termites are social insects
that are able to feed on wood because of a community of protists and
bacteria that live in the insects' guts. The microorganisms produce the enzymes which digest the cellulose in the wood.
Cleaning symbiosis. (Source: Sodwana Diving)
2.
Defensive mutualisms are interactions in which one species
receives food or shelter in return for protecting its partner species
from predators or parasites. Some species provide direct food rewards to
a species whose presence decreases risk of
predation. Two well-known examples involving ants are associations between
Pseudomyrmex ants and
Acacia shrubs in Central America, and between wood ants and aphids in the
forests
of Europe and North America. In cleaning symbioses, a cleaner species
receives a food reward by removing (and eating) parasites from the body
of its partner. Many biologists consider some forms of mimicry to be
special cases of defensive mutualism. For example, the aposematic
species involved in a Mullerian mimetic association all converge on the
same coloration, thereby reducing the number of patterns that local
predators have to learn. All the prey species involved benefit because
predators avoid them.
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